Reise, H. Schwarzer, A.-K., Hutchinson, J.M.C. & Schlitt, B. 2020. Genital morphology differentiates three subspecies of the terrestrial slug Arion ater (Linnæus, 1758) s.l. and reveals a continuum of intermediates with the invasive A. vulgaris Moquin-Tandon, 1855. Folia Malacologica 28: 1–34.

Doi: https://doi.org/10.12657/folmal.028.001 (downloads free, maybe after free registration).

Or, pdf (25 MB) downloadable from here.
Link to the article in the journal (downloads free after free registration).

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