J.M.C. HUTCHINSON 1990. Stabilising selection in Weldons snails: a reappraisal.
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DOI https://doi.org/10.1038/hdy.1990.14 to access article via journal (free).
I overlooked another paper that applied a similar approach with scale number in lizards: Cook, L.M. 1979. Variation in the Madeiran lizard, Lacerta dugesii. Journal of Zoology 187: 327340.
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